TMNT 2012: Complete Lore Teenage Mutant Ninja Turtles (2012) follows four mutant turtle brothers raised beneath New York City by Master Splinter, formerly Hamato Yoshi. Trained in ninjutsu, the brothers defend their home against mutants, aliens, criminals, and their father's greatest enemy, the Shredder. Origins: Hamato Yoshi belonged to Japan's Hamato Clan alongside Oroku Saki, his adopted brother and eventual rival. Both loved Tang Shen, who chose Yoshi. Consumed by jealousy, Saki attacked their home, causing Shen's death. He kidnapped Yoshi's daughter Miwa, renamed her Karai, and raised her believing Yoshi was responsible. Years later, Yoshi encountered four baby turtles and mysterious aliens called the Kraang. Exposure to mutagen transformed Yoshi into a humanoid rat and the turtles into humanoid mutants. He adopted and trained them. The Brothers: - Leonardo (Blue): Disciplined leader wielding twin katanas. Responsible, honorable, protective, and occasionally insecure. - Raphael (Red): Hotheaded, sarcastic, fiercely loyal fighter wielding twin sai. Often clashes with Leonardo. - Donatello (Purple): Brilliant scientist and inventor wielding a bo staff. Has feelings for April O'Neil. - Michelangelo (Orange): Playful, compassionate, unpredictable youngest brother wielding nunchaku. Season 1: The turtles discover the surface world and rescue April O'Neil from the Kraang, aliens planning to transform Earth. April becomes their friend and Splinter's student. Meanwhile, Shredder discovers Yoshi survived and sends his Foot Clan after the turtles. Leonardo encounters Karai, unaware she is Splinter's daughter. The turtles battle mutants, Foot ninjas, and Kraang technology before defeating the Kraang's initial invasion. Season 2: Mutagen spreads throughout New York, creating dangerous mutants. The turtles meet Casey Jones, a hockey-masked vigilante, and confront Tiger Claw, Rahzar, Slash, and other enemies. April discovers her psychic powers originate from Kraang experimentation. Karai learns Splinter is her real father but is eventually mutated into a serpent. The Kraang invade New York, Leonardo is badly injured, and the brothers retreat to April's farmhouse. Season 3: Leonardo recovers while the brothers encounter supernatural threats. Returning to New York, they reunite with Splinter and fight to reclaim their city. They gain allies including the Mighty Mutanimals and confront Bebop and Rocksteady. Eventually, the alien Triceratons arrive, intending to destroy Earth. Shredder attacks Splinter as he tries to stop their black-hole generator. Earth is destroyed, but the robotic Fugitoid rescues the turtles, April, and Casey, transporting them into the past. Season 4: Traveling through space, the heroes battle alien enemies and attempt to prevent Earth's destruction. Raphael meets Mona Lisa, a Salamandrian warrior. After saving Earth, they return to New York. Shredder deliberately mutates himself into Super Shredder, becoming monstrously powerful. He eventually kills Splinter, devastating the brothers. Leonardo leads his family into a final confrontation and defeats Super Shredder, ending their father's lifelong feud. Season 5: The turtles continue protecting New York without Splinter. Tiger Claw attempts to resurrect Shredder using the demon Kavaxas. Other adventures feature monsters, supernatural enemies, and a crossover with the 1987 turtles. The alternate-future Mutant Apocalypse storyline depicts the brothers surviving a ruined world. Legacy: April becomes a powerful kunoichi, Casey remains a loyal ally, and Karai reclaims her Hamato heritage. Leonardo grows into a true leader, Raphael learns restraint, Donatello develops confidence, and Michelangelo proves his compassion is a strength. Despite loss, betrayal, and impossible battles, the brothers preserve Splinter's teachings. Together, they are the Teenage Mutant Ninja Turtles, protectors of New York and heirs to the Hamato Clan.