The leader of the 'Zodiac Alliance,' an organization that has awakened from a 100-year slumber to resume the battle against the Mysteries. Representing the Cat of the Zodiac Alliance, he transforms into Kamen Rider Mao using a special belt. He possesses the charisma to gather the alliance members, who are all eccentric individuals, but he also has a whimsical side. He boasts remarkable strength even among the Zodiac Riders. Transformation: Kamen Rider Mao (Cat) Transformation Sound: [Baku! En! Retsu! Go!] [Born! Kamen Rider Mao!]

A young man who got caught up in the battle between Mice and the Mysteries and met Sora's group. In reality, he is a member of the 'Zodiac Alliance' trying to uncover the secrets of Sora and Mice. Ordered by Kamuri to conduct an undercover investigation at the Mizuki residence, he became Yuna's exclusive tutor. He has a serious personality that clashes with Sora, but they occasionally display excellent teamwork when their interests align. Transformation: Kamen Rider Read (Cat) Transformation Sound: [Myo! Kamen Rider Read!]

The Rabbit of the Zodiac Alliance. He transforms into Kamen Rider Dat. He has excellent hearing and can detect the appearance of Mysteries. Confident in his speed, he is always timing his own records with a pocket watch. He is devoted to Kamuri. He is a man of few words and has a blunt personality, with a temper so short that his words and actions are always rushed. Transformation: Kamen Rider Dat (Rabbit) Transformation Sound: [Myo! Kamen Rider Dat!]

The Snake of the Zodiac Alliance. He transforms into Kamen Rider Jao. Boasting high combat power, he overwhelms Mysteries with continuous attacks from the longest tail among the Zodiac and his scythe. He believes that might is right and has sworn loyalty to Kamuri. Transformation: Kamen Rider Jao (Snake) Transformation Sound: [Sa! Kamen Rider Jao!]

A veteran of the Zodiac Alliance. He has more experience fighting Mysteries than anyone else and is highly regarded even by Kamuri. Transformation: Kamen Rider Ryuzu (Dragon) Transformation Sound: [Jin! Kamen Rider Ryuzu!]

The Sheep of the Zodiac Alliance. He transforms into Kamen Rider Muton. Usually, he does nothing but sleep, but when he thinks of Banken, whom he calls 'Ken-nim,' he displays a level of 'affection' that even the other Zodiac members cannot handle. Transformation: Kamen Rider Muton (Sheep) Transformation Sound: [Mi! Kamen Rider Muton!]

The Rooster of the Zodiac Alliance. He transforms into Kamen Rider Kei. The youngest in the Zodiac Alliance. He is quickly adapting to modern civilization, such as being proficient with smartphones. Transformation: Kamen Rider Kei (Rooster) Transformation Sound: [Yu! Kamen Rider Kei!]

The Dog of the Zodiac Alliance. He transforms into Kamen Rider Banken. A literary man with deep knowledge of poetry, always carrying a book of poems. He uses his keen sense of smell to quickly locate and confront Mysteries. Transformation: Kamen Rider Banken (Dog) Transformation Sound: [Sul! Kamen Rider Banken!]

The Tiger of the Zodiac Alliance. He transforms into Kamen Rider Tigre. He has excellent insight, a brotherly personality, and is calm and composed. While reliable, he can show a harsh side during life-or-death battles with Mysteries. He is also a long-time friend who has relied on the leader, Kamuri, for years. Transformation: Kamen Rider Tigre (Tiger) Transformation Sound: [In! Kamen Rider Tigre!]